The pair - who played Hermione Granger and Ron Weasley in the wizard franchise - topped the list compiled by Forbes after banking $4.2 billion between them at the global box office in four movies in the series in the last five years.
`Twilight` actors Robert Pattinson and Kristen Stewart - who are dating each other in real life - came second after they picked up $2.5 billion from their portrayals of vampire Edward Cullen and human Bella Swan respectively in the saga.
Shia LaBeouf and Megan Fox earned $1.5 billion after they starred as onscreen lovers Sam Witwicky and Mikaela Banes in `Transformers` and `Transformers: Revenge of the Fallen`, which narrowly beat the actor`s onscreen romance with Rosie Huntington-Whiteley in `Transformers: Dark of the Moon`, which pulled in $1.1 billion.
Robert Downey Jr. and Gwyneth Paltrow`s portrayals of Tony Stark and Pepper Potts in the `Iron Man` films saw them claim fourth with $1.2 billion.
Among the other stars who made up the list were Johnny Depp and Penelope Cruz, who earned $1 billion for `Pirates of the Caribbean: On Stranger Tides`, ahead of Orlando Bloom and Keira Knightley`s $963 million for their romance in the first three movies of the series.
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